Making some progress. 'Multilevel Selection Processes in Economics: Theory and Methods' by Natalia Zinovyeva (http://www3.druid.dk/wp/20040008.pdf )has a little example which helped. Another paper 'Understanding microbial cooperation' by James Damore and Jeff Gore (Supplement http://scholar.harvard.edu/files/james/files/damore_jtb_2012_supp.pdf?m=1363388631, to orinial paper http://www.ncbi.nlm.nih.gov/pmc/articles/PMC3176967/)has a fairly comprehensive discussion and also a discussion of some of the difficulties.
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